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Cynipoidea of the world

(Life: Kingdom: Metazoa (animals); Phylum: Arthropoda; Class: Hexapoda; Order: Hymenoptera)


Identification keys






Distribution Worldwide.
Biology Parasitoids or phytophages (gall formers and inquilines).


Abe, Y., T. Ide, and K.-I. Odagiri. 2014. Cynipidae (Hymenoptera: Cynipoidea) on Cyclobalanopsis (Fagaceae) in mainland China, with the first record of sexual generation of Aguiar AP, Deans AR, Engel MS, Forshage M, Huber JT, Jennings JT, Johnson NF, Lelej AS, Longino JT, Lohrmann V, Miko I. 2013. Order Hymenoptera. In: Zhang, Z.-Q.(Ed.) Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013). Zootaxa 3703: 51-62. http://dx.doi.org/10.11646/zootaxa.3703.1.12

Aluja, M.; Lopez, M.; Sivinski, J. 1998. Ecological evidence for diapause in four native and one exotic species of larval-pupal fruit fly (Diptera: Tephritidae) parasitoids in tropical environments Annals of the Entomological Society of America 91:821-833

Ashmead, W.H. 1900. Notes on some New Zealand and Australian Parasitic Hymenoptera, with descriptions of new genera and new species. Proceedings of the Linnean Society of New South Wales 25: 327–360.

Bassett, H. F. 1881. New Species of Cynipidae. The Canadian Entomologist. 13: 51–53. doi:10.4039/Ent1351-3

Benoit, P.L.G. 1955. Les Liopteridae Oberthurellinae du Congo Belge. Revue de Zoologie et de Botanique Africaines 52: 283–293.

Benoit, P.L.G. 1956. Liopteridae-Mesocynipinae nouveaux du Congo Belge (Hym.-Cynipoidea). Revue de Zoologie et de Botanique Africaines 53: 51-56.

Borgmeier, T. 1935. Sobre alguns Cynipideos parasiticos e cecidogenos do Brasil (Hymenoptera, Cynipidae). Arquivos do Instituto de Biologia Vegetal 2: 97-124.

Branstetter MG, Childers AK, Cox-Foster D, Hopper KR, Kapheim KM, Toth AL, Worley KC. 2018. Genomes of the Hymenoptera. Current Opinion in Insect Science 25: 65-75. https://doi.org/10.1016/j.cois.2017.11.008

Buffington, M. L. 2007. The occurrence and phylogenetic implications of the ovipositor clip within the Figitidae (Insecta: Hymenoptera: Cynipoidea). J. Nat. His. 41: 2267–2282.

Buffington, M.L. 2011. Description, circumscription and phylogenetics of the Diglyphosematini Belizin, 1961, and the description of a new genus (Hymenoptera: Figitidae: Eucoilinae). Proc. Entomol. Soc. Wash. 113: 239–290.

Buffington, M.L. 2012. Description of Nanocthulhu lovecrafti, a preternatural new genus and species of Trichoplastini (Figitidae: Eucoilinae). Proceedings of the Entomological Society of Washington 114(1): 5-15.

Buffington, ML & Copeland, R.S.  2015. Muhaka icipe, an enigmatic new genus and species of Kleidotomini (Hymenoptera: Figitidae: Eucoilinae) from an East African coastal forest. Journal of Natural History 49: 1-11. DOI: 10.1080/00222933.2015.1042411

Buffington, M. L., and J. Liljeblad 2008. The description of Euceroptrinae, a new subfamily of Figitidae (Hymenoptera), including a revision of Euceroptres Ashmead, 1896, and the description of a new species. Journal of Hymenoptera Research 17: 44–56.

Buffington, M.L. & van Noort, S. 2007. A world revision of Pycnostigminae (Cynipoidea: Figitidae) with descriptions of seven new species. Zootaxa 1392: 1-30

Buffington, M.L. & van Noort, S. 2009. A world revision of Anacharoides Cameron, 1904 (Hymenoptera: Figitidae) with a description of a new species. Zookeys 20: 245-274.

Buffington, M.L. & van Noort, S. 2012. Revision of the Afrotropical Oberthuerellinae (Cynipoidea: Liopteridae). Zookeys 202: 1-154.

Buffington, M. L., S. Brady, S. Morita, and S. van Noort. 2012. Divergence estimates and early evolutionary history of Figitidae (Hymenoptera: Cynipoidea). Systematic Entomology 37: 287–304.
Buffington ML, Forshage M, Liljeblad J, Tang CT, van Noort, S. 2020. World Cynipoidea (Hymenoptera):a key to higher-level groups. Insect Systematics and Diversity 4(4): 1–69. https://doi.org/10.1093/isd/ixaa003

Buffington ML, Giorgini M, Lue C-H, Formisano G, Cascone P, Forshage M, Driskell A & Guerrieri E. 2020. Description of the aberrant Leptopilina lasallei n. sp., with an updated phylogeny of Leptopilina Förster (Hymenoptera: Figitidae: Eucoilinae). Journal of Natural History 54: 565-583. https://doi.org/10.1080/00222933.2020.1754483

Burks, B.D. 1979. Superfamily Cynipoidea, pp. 1045- 1107. In K. V. Krombein, P. D. Hurd, Jr., D. R. Smith, and B. D. Burks (eds), Catalog of Hymenoptera in America North of Mexico. Volume 1, Smithsonian Institution Press, Washington, DC.

Cameron, P. 1904. Descriptions of new genera and species of Hymenoptera from Dunbrody, Cape Colony. Records of the Albany Museum 1: 125-160.

Cameron, P. 1905. A new genus and species of Cynipidae from South Africa, representing a new subfamily. Annals and Magazines of Natural History, Series 7, 16,  20-21.

Chai M, Liu Z, Niu Y, Dong Y & Wang Y. 2018. Discovery of two new species of the genus Heteribalia Sakagami (Hymenoptera: Ibaliidae) from China. Entomotaxonomia 40: 217-230. 10.11680/entomotax.2018024.

Cuesta-Porta, V., Arnedo, M.À., Cibrián-Tovar, D., Barrera-Ruiz, U.M., García-Martiñón, R.D., Equihua-Martínez, A., Estrada-Venegas, E.G., Clark-Tapia, R., Romero-Rangel, S. and Pujade-Villar, J., 2020. A new genus of oak gall wasp, Striatoandricus Pujade-Villar (Hymenoptera: Cynipidae: Cynipini) from America with descriptions of two new Mexican species. Zoological Studies 59: 8 (2020). https://doi.org/10.6620/ZS.2020.59-08

Benoit, P.L.G. 1956. Figitidae - Aspicerinae nouveaux du Congo Belge. Revue Zool. Bot. afr. 53: 195-204.

Diaz, N. 1977. Estudio ecológico y sistemático de Cinipoideos Neotropicales II (Hymenoptera). Epicoela rubicunda Borgmeier y E. seminigra n. sp. Neotrópica 23(70): 189-192.

Diaz, N. & F. Gallardo. 1996. Sobre cinipoideos del Brasil, parasitoides de dipteros estercoleros (Hymenoptera: Cynipoidea). Rev. Soc. Entomol. Argent. 55: 127-129.

Diaz, N.B., Gallardo, F.E., Marchiori, C.H., & Linhares, A.X. 2000. Cynipoidea parasitoids of dung-flies in Brazil. II (Insecta: Hymenoptera). Anais da Sociedade Entomológica do Brasil 29(3): 469-474.

Díaz, N.B., Gallardo, F.E., Gaddi, A.L., & Walsh, G.C. 2009. Description of a new genus and species of Eucoilinae (Hymenoptera: Cynipoidea: Figitidae) parasitoid of Ephydridae (Diptera). Annals of the Entomological Society of America, 102(4): 603-607.

Fergusson, N. 1992. A remarkable new genus and species of Cynipoidea (Hymenoptera) from Papua New Guinea. Journal of Natural History 26(3): 659-662.

Ferrer-Suay, M., Selfa, J. & Pujade-Villar, J. 2012. First record of Alloxysta Förster from Madagascar, with descriptions of two new species (Hymenoptera: Cynipoidea: Figitidae: Charipinae). African Entomology 20: 222-228.

Ferrer-Suay, M., Selfa, J., Pujade-Villar, J. 2013. A review of Alloxysta species (Hymenoptera: Cynipoidea: Figitidae: Charipinae) from Africa. African Entomology 21: 255266.

Ferrer-Suay, M., Paretas-Martinez, J., & Pujade-Villar, J. 2013. Revision of Apocharips Fergusson (Hymenoptera: Figitidae: Charipinae) with description of three new species from Colombia. Zootaxa, 3646(4), 487–500. http://dx.doi.org/10.11646/zootaxa.3646.4.8

Ferrer-Suay, M, Carpenter, JM, Lebeau, C, Pujade-Villar J. 2017. Designation of lectotypes for the Mexican species of Andricus described by Alfred Kinsey and comments about some generic synonymies (Hymenoptera: Cynipidae). Entomologica Americana 123: 29-34. https://doi.org/10.1664/1947-5144-123.1-4.29
Foelker, C. J., Standley, C. R., Fierke, M. K., Parry, D. and Whipps, C. M. 2015. Host tissue identification for cryptic hymenopteran parasitoids associated with Sirex noctilio. Agricultural and Forest Entomology. doi: 10.1111/afe.12137

Fontal-Cazalla, F. M., Buffington, M. L., Nordlander, G., Liljeblad, J., Ros-Farré, P., Nieves-Aldrey, J. L., Pujade-Villar, J. & Ronquist, F. (2002). Phylogeny of the Eucoilinae (Hymenoptera: Cynipoidea: Figitidae). Cladistics 18, 154–199.

Forshage, M & Nordlander, G. 2018. The identity of figitid parasitoids (Hymenoptera: Cynipoidea: Figitidae) of anthomyiid flies in conifer cones. European Journal of Entomology 115: 104-111.

Gillette, C.P. 1891. Descriptions of new Cynipidae in the collection of the Illinois State Laboratory of Natural History. Bulletin of the Illinois State Laboratory of Natural History 3: 191-197.

Guimaraes, J. A.; Zucchi, R. A.; Diaz, N. B.; de Souza Filho, M. F.; Uchoa F., M. A. 1999. Espzcies de Eucoilinae (Hymenoptera: Cynipoidea: Figitidae) Parasitides de Lavas Frug’voras (Diptera: Tephritidae e Lonchaeidae) no Brasil. Anais da Sociedade Entomologica do Brasil 28:263-273.

Hedicke, H. 1930. Beiträge zur Kenntnis der Cynipiden. (Hym.) XVI Zur Synonymie der Anacharitinen, Eucoilinen und Figitinen. Deutsche Entomologische Zeitschrift 1930: 74-76

Hedicke, H., and G. Kerrich. 1940. A revision of the family Liopteridae (Hymenopt., Cynipoidea). Transactions of the Royal Entomologocal Society, London 90: 177–228.

Ide T, Mu MA, Nobuyuki T. 2020. First record of Cynipidae from Myanmar with description of a new species of Lithosaphonecrus (Hymenoptera: Cynipidae: Synergini). Zootaxa 4810(2): 344–350. https://doi.org/10.11646/zootaxa.4810.2.7

Julien, M.H. & Control, A. 2014. Biological Control of Weeds: A World Catalogue of Agents and Their Target Weeds. XF2006287138.

Johnson BR, Borowiec ML, Chiu JC, Lee EK, Atallah J, Ward PS. 2013. Phylogenomics resolves evolutionary relationships among ants, bees, and wasps. Current Biology 23: 2058–2062. https://doi.org/10.1016/j.cub.2013.08.050

Kieffer, J.J. 1904. Les Cynipides. Supplement aux Cynipides. Eucoelinae. Hym. Hym. Eur. 7: 603–631.

Liljeblad, J., Nieves-Aldrey, J.L., Neser, S. & Melika, G. 2011. Adding another piece to the cynipoid puzzle: the description of a new tribe, genus and species of gall wasp (Hymenoptera: Cynipidae) endemic to The Republic of South Africa. Zootaxa 2806: 35–52.

Lobato-Vila, I., Pujade-Villar, J. 2019. Revision of world Ceroptresini (Hymenoptera: Cynipidae) with the description of a new genus and five new species. Zootaxa. 4685: 1–67. http://dx.doi.org/10.11646/zootaxa.4685.1.1

Liu, Z. 1998. A new species of Ibalia from Borneo, and revised phylogeny and historical biogeography of Ibaliidae (Hymenoptera: Cynipoidea). Journal of Hymenoptera Research 7, 149–156. https://doi.org/10.5962/bhl.part.29275

Liu, Z. 2009. Phylogeny, Biogeography, and Revision of the Subfamily Dallatorrellinae (Hymenoptera: Liopteridae). American Museum Novitates. 3353. 1-23. https://doi.org/10.1111/j.1365-3113.1995.tb00099.x

Liu Z & Nordlander G. 1994. Review of the family Ibaliidae (Hymenoptera: Cynipoidea) with keys to the genera and species of the World. Insect Systematics & Evolution 24 (6): 377–392.  https://doi.org/10.1163/187631294X00153

Liu, Z, Ronquist, F. & Nordlander, G. 2007. The cynipoid genus Paramblynotus: revision, phylogeny, and historical biogeography (Hymenoptera, Liopteridae). Bulletin of the American Museum of Natural History 304: 1-151.

Lue, C. H., A. Driskell, J. Leips, and M. L. Buffington. 2016. Review of the genus Leptopilina (Hymenoptera, Cynipoidea, Figitidae, Eucoilinae) from the Eastern United States, including three newly described species. J. Hym. Res. 53: 35–76.

Mata-Casanova, N., Pujade-Villar, J. 2013. Acanthaegilopsis malagasy gen. n. and sp. n. of Anacharitinae (Hymenoptera: Cynipoidea: Figitidae) from Madagascar and the Comoros. African Entomology 21(1): 161–164.

Mata-Casanova N, Pujade-Villar J. 2013. Nuevas aportaciones al conocimiento de Acanthaegilips en Colombia (Hymenoptera: Figitidae: Anacharitinae). Revista colombiana de Entomología 39(1):71–75.

Mata-Casanova N, Selfa J, Arcaya E, Sosa F, Tormos J, Pujade-Villar J. 2014. First host record for Acanthaegilips (Hymenoptera: Figitidae: Anacharitinae) and description of a new species from Venezuela. Fla Entomol 97(2):461–464.

Mata-Casanova N,  Selfa J  & Pujade-Villar J. 2014. Description of two new species of Anacharitinae (Hymenoptera: Figitidae) from the Afrotropical region. African Entomology 22: 545-551. doi: http://dx.doi.org/10.4001/003.022.0312

Mata-Casanova N, Selfa J & Pujade-Villar J. 2015. Description of two new species of Anacharis Dalman, 1823 (Hymenoptera: Figitidae, Anacharitinae) from the Afrotropical Region. African Entomology 23: 314–320. http://dx.doi.org/10.4001/003.023.0222

Mata-Casanova N, Selfa J & Pujade-Villar J. 2017. Revision of the current knowledge of genus Aegilips Haliday, 1835 (Hymenoptera: Figitidae: Anacharitinae) in the Afrotropical Region, with description of three new species. African Entomology 25: 62 – 67. http://dx.doi.org/10.4001/003.025.0062

Mata-Casanova, N., Selfa, J., Pujade-Villar, J. 2018. The true identity of Amphithectus austriacus (Tavares, 1928): junior synonym of Amphithectus opacus (Thomson, 1862) n. comb. Butlletí de la Institució Catalana d'Història Natural 82: 137-139.

Mayr, G. 1881. Die genera der gallenbewohnenden Cynipiden. Jahresberichte der Rossauer Communal Oberrealschule im I. Bezirke 20:138.

Melika, G. 2006. Gall wasps of Ukraine. Cynipidae. Vestnik zoologii, supplement 21, vol. 1–2.
Melika, G. 2012. Superfamily Cynipoidea – cynipoids or gallwasps, pp. 139–146. In A. Lelej (ed.) Annotated catalogue of the insects of the Russian Far East. Vol. I. Hymenoptera. Vladivostok. Dalnauka, Russia (in Russian)
Melika, G., and W. G. Abrahamson. 2002. Review of the world genera of oak cynipid wasps (Hymenoptera: Cynipidae: Cynipini), pp. 150–190. In G. Melika, and C. Thuroczy (eds.), Parasitic Wasps: evolution, systematics, biodiversity and biological control. Agroinform, Budapest, Hungary.

Melika, G. & Nicholls, J.A. 2021. A new genus of Nearctic oak gall wasp, Grahamstoneia Melika & Nicholls, gen. nov. (Hymenoptera: Cynipidae, Cynipini). Zootaxa 4999(5): 456–468. https://doi.org/10.11646/zootaxa.4999.5.4

Melika, G., J. Pujade-Villar, Y. Abe, C. T. Tang, J. A. Nicholls, N. Wachi, T. Ide, M. M. Yang, Z. Pénzes, G. Csóka, and G. N. Stone. 2010. Palaearctic oak gallwasps galling oaks (Quercus) in the section Cerris: re-appraisal of generic limits, with descriptions of new genera and species (Hymenoptera: Cynipidae: Cynipini). Zootaxa. 2470: 1–79.
Melika, G., C. T. Tang, J. A. Nicholls, M. M. Yang, and G. N. Stone. 2011. Four new species of Dryocosmus gallwasps from Taiwan (Hymenoptera: Cynipidae: Cynipini). ISRN Zoology. 17 pp. doi:10.5402/2011/725180
Melika, G., E. Buss, J. Nicholls, J. Platt Bird, and G. N. Stone. 2013. Life cycle of Discholcaspis quercusvirens (Hymenoptera: Cynipidae) with a description of the sexual generation. Florida Entomol. 96: 991–1001.
Melika, G., C. T. Tang, F. Sinclair, M. M. Yang, K. Lohse, J. Hearn, J. A. Nicholls and G. N. Stone. 2013. A new genus of oak gallwasp, Cyclocynips Melika, Tang & Sinclair (Hymenoptera: Cynipidae: Cynipini), with descriptions of two new species from Taiwan. Zootaxa.
3630: 534–548.

Melika G, Pujade-Villar J, Nicholls JA, Cuesta-Porta V, Cooke-Mcewen C, Stone GN. 2021. Three new Nearctic genera of oak cynipid gall wasps (Hymenoptera: Cynipidae: Cynipini): Burnettweldia Pujade-Villar, Melika & Nicholls, Nichollsiella Melika, Pujade-Villar & Stone, Disholandricus Melika, Pujade-Villar & Nicholls and re-establishment of the genus Paracraspis Weld. Zootaxa 4993(1): 1–81. https://doi.org/10.11646/zootaxa.4993.1.1

Nielsen, M. & Buffington, M. 2011. Redescription of Stentorceps Quinlan, 1984 (Hymenoptera: Figitidae), with a description of five new species. African Entomology 19: 597–613.

Nieves-Aldrey, J.L. 1994. Revision of the West-European genera of the tribe Aylacini Ashmead (Hymenoptera, Cynipidae). Journal of Hymenoptera Research 3: 175–206.

Nieves-Aldrey, J. L. 2001. Hymenoptera: Cynipidae: Fauna Iberica, vol. 16, 636 pp. M. A. Ramos et al. (eds.). Museo Nacional de Ciencias Naturales. CSIC, Madrid, Spain.
Nieves-Aldrey, J. L. 2002. Notes on the morphology and biology of Neaylax salviae (Giraud), a cynipid new to the Iberian fauna, with a description of the larva (Hymenoptera, Cynipidae). Bol. Soc. Entomol. Arag. 31: 61–65.
Nieves-Aldrey, J. L. 2012. Revision of the Amphibolips species of Mexico excluding the “niger complex” Kinsey (Hymenoptera: Cynipidae), with description of seven new species. Zootaxa 3545: 1–40.
Nieves-Aldrey, J. L., J. Liljeblad, M. Hernández-Nieves, A. Grez, and J. A. A. Nylander. 2009. Revision and phylogenetics of the genus Paraulax Kieffer (Hymenoptera, Cynipidae) with biological notes and description of a new tribe, a new genus, and five new species. Zootaxa. 2200: 1–40.

Nieves-Aldrey JL, Blas GS. 2015. Revision of the Neotropical genus Eschatocerus Mayr (Hymenoptera, Cynipidae, Eschatocerini) with biological notes and the first description of the terminal larva. Zootaxa 4012: 135-155. https://doi.org/10.11646/zootaxa.4012.1.7

Nieves-Aldrey J.L., J. Liljeblad, M. Hernández-Nieves, A Grez and J.A.A. Nylander. 2009. Revision and phylogenetics of the genus Paraulax Kieffer (Hymenoptera, Cynipidae) with biological notes and description of a new tribe, a new genus, and five new species Zootaxa 2200: 1-40. 10.5281/zenodo.189597.

Nordlander, G. 1982. Identities and relationships of the previously confused genera Odonteucoila, Coneucoela, and Trichoplasta (Hymenoptera, Cynipoidea: Eucoilidae). Insect Systematics & Evolution 13: 269–292. https://doi.org/10.1163/187631282X00327

Ovruski, S.; Aluja, M.; Sivinski, J.; Wharton, R. 2000. Hymenopteran parasitoids on fruit-infesting Tephritidae (Diptera) in Latin America and the southern United States: Diversity, distribution, taxonomic status and their use in fruit fly biological control. Integrated Pest Management Reviews 5:81-107.

Paretas-Martinez, J.; Melika, G.; Pujade-Villar, J. 2009. Description of four new species of Dilyta Förster (Hymenoptera: Figitidae: Charipinae) from the Afrotropical Region. African Entomology 17: 207-214.

Paretas-Martínez, J., Arnedo, M.A., Melika, G., Selfa, J., Seco-Fernández, M.V., Fülöp, D., Pujade-Villar, J. 2007. Phylogeny of the parasitic wasp subfamily Charipinae (Hymenoptera, Cynipoidea, Figitidae) Zoologica Scripta 36: 153–172.

Paretas-Martínez, J., Restrepo-Ortiz C., Buffington, M. L. and Pujade-Villar, J. 2011. Systematics of Australian Thrasorinae (Hymenoptera, Cynipoidea, Figitidae) with descriptions of Mikeiinae, new subfamily, two new genera, and three new species. ZooKeys 108: 21-48. 10.3897/zookeys.108.829. https://doi.org/10.3897/zookeys.108.829

Peters RS, Krogmann L, Mayer C, Donath A, Gunkel S, Meusemann K, Kozlov A, Podsiadlowski L, Petersen M, Lanfear R, Diez PA. 2017. Evolutionary history of the Hymenoptera. Current Biology 27:1013-1018. https://doi.org/10.1016/j.cub.2017.01.027

Pujade-Villar, J. 2019. Reclasificación de los grandes taxa de Cynipoidea (Hymenoptera) desde Weld (1952) hasta 2017: la historia de una lucha evolutiva entre el parasitoidismo y la fitofàgia. Folia Entomol. Mex. 5: 18−65.

Pujade-Villar, J., and M. Ferrer-Suay. 2015. Adjudicació genèrica d’espècies mexicanes d’ubicació dubtosa descrites per Kinsey i comentaris sobre la fauna mexicana (Hymenoptera: Cynipidae: Cynipini). Butll. Inst. Catalana Hist. Nat. 79: 7–14.
Pujade-Villar, J., and G. Melika. 2014. Re-establishment of Erythres Kinsey, 1937 as a valid genus of gallwasps from Mexico (Hymenoptera: Cynipidae: Cynipini). Dugesiana. 21: 155–160.
Pujade-Villar, J., Ros-Farré, P. & Brandao, C.R.F. 2000. Notes on Brazilian Liopteridae (Hymenoptera: Cynipoidea), with the description of Pseudibalia angelicae n. sp. Papéis avulsos de zoologia (Brasil) 41(19): 289-301.

Pujade-Villar, J. & Rodriguez, P. 2015. Primera cita del género Melikaiella (Hym.: Cynipidae) para Colombia y descripción de una especie nueva. Orsis 29. 193. 10.5565/rev/orsis.23.

Pujade-Villar, J. & Schiopu, I. 2015. Species of Aylacini Ashmead, 1903 (Hymenoptera: Cynipidae) inducing galls on Papaver L. (Papavaceae) from Romania with description of a new species and notes on parasitoids (Hymenoptera). Zoosystema 37: 505-515. https://doi.org/10.5252/z2015n3a5

Pujade-Villar, J., D. Bellido, G. Segú, and G. Melika. 2001. Current state of knowledge of heterogony in Cynipidae (Hymenoptera, Cynipoidea). Sessió Conjunta de Entomologia ICHN-SCL. 11: 87–107.

Pujade-Villar, J., S. Romero-Rangel, C. Chagoyán-García, A. Equihua-Martínez, E. G. Estrada-Venegas, and G. Melika. 2010. A new genus of oak gallwasps, Kinseyella Pujade-Villar & Melika, with a description of a new species from Mexico (Hymenoptera: Cynipidae: Cynipini). Zootaxa. 2335: 16–28.

Pujade-Villar, J., Hanson, P., Medina, C. A., Torres, M., and Melika, G. 2012. A new genus of oak gallwasps, Zapatella Pujade-Villar & Melika, gen. n., with a description of two new species from the Neotropics (Hymenoptera, Cynipidae, Cynipini). ZooKeys 210: 75-104.

Pujade-Villar, J., P. Hanson, and G. Melika. 2012. A new genus of oak gallwasp, Coffeikokkos Pujade-Villar & Melika, gen. n., with a description of a new species from Costa Rica (Hymenoptera, Cynipidae). ZooKeys. 168: 19–29.

Pujade-Villar, J., A. Equihua-Martínez, E. G. Estrada-Venegas, and G. Melika. 2013. A new genus of oak gallwasp, Kokkocynips Pujade-Villar & Melika gen. n., with a description of a new species from Mexico (Hymenoptera: Cynipidae). Acta Zool. Mexicana. 29: 209–218.

Pujade-Villar, J., Cibrián-Tovar, D., Cibrián-Llanderal, V. D., Equihua-Martínez, A., Estrada-Venegas, E. G., Serrano-Muñoz, M. and Lomeli-Flores, J. R. 2014. A new genus of oak gallwasp, Melikaiella Pujade-Villar (Hymenoptera: Cynipidae: Cynipini), from the Nearctic region. Dugesiana 21: 1-29.

Pujade-Villar, J., I. Lobato-Vila, and M. Ferrer-Suay. 2017. Restablecimiento del género Dros Kinsey (Hymenoptera: Cynipidae: Cynipini) como género válido para especies Americanas. Entomol. Mexicana. 4: 752–758.

Pujade-Villar, J., Wang, Y. & Lobato-Vila, I. 2020. Lithosaphonecrus puigdemonti sp. nov. from China (Hymenoptera: Cynipidae). Entomotaxonomia, 42 (1), 70–80.

Pujade-Villar J, Restrepo-Ortiz CX, García JL, Ros-Farré P. 2009. Redescripción de Acanthaegilips Ashmead 1897 y descripción de dos nuevas especies (Figitidae: Anacharitinae). Dugesiana 16(2):57–65
Pujade-Villar J, Restrepo-Ortiz CX, Ros-Farré P, García JL. 2009. Acanthaegilips Ashmead, 1897 (Cynipoidea, Figitidae: Anacharitinae) de Venezuela. Entomotropica 24(1):23–30.

Pujade-Villar, J., Kang, M., Bae, J., Wang, Y.-P., Guo, R. & Jung, S. 2020. Additional records to the Korean Cynipini: subspecies description, new combinations and checklist (Hymenoptera, Cynipidae). Journal of Asia-Pacific Entomology 23: 1208–1221.
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Photographs © Simon van Noort (Iziko Museums of South Africa), or

Photographs © Matt Buffington (USDA/ARS).

Matt Buffington (Systematic Entomology Laboratory, USDA/ARS) and Mattias Forshage (Swedish Museum of Natural History) both played an integral role in the development of the Cynipoidea pages.

Web author Simon van Noort (Iziko South African Museum)


Citation: van Noort, S. 2024. WaspWeb: Hymenoptera of the World. URL: www.waspweb.org (accessed on <day/month/year>).

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