Coelioxys - WaspWeb

Coelioxys afra Lepeletier, 1941 (Democratic Republic of the Congo, Eritrea, Gambia, Malawi, Mozambique, Nigeria, South Africa, Togo. Also in the Palaearctic region)

Coelioxys afra melanafra Cockerell, 1935 (South Africa)

Coelioxys albociliata Pasteels, 1968 (Guinea)

Coelioxys albomarginata Friese, 1922 (Democratic Republic of the Congo, Tanzania)

Coelioxys analis Friese, 1911 (Malawi, South Africa, Zambia, Zimbabwe)

Coelioxys apicata Smith, 1854 (Gabon, Ghana, Mozambique, Senegal, Sierra Leone, Zambia) Also in Oriental region)

Coelioxys argentipes Smith, 1879 (Malawi, Sierra Leone)

Coelioxys ateneata Strand, 1921 (Democratic Republic of the Congo, Mali, Senegal, South Africa, Uganda)

Coelioxys aureociliata Pasteels, 1968 (Senegal)

Coelioxys aurifrons Smith, 1853 (Congo, Democratic Republic of the Congo, Ethiopia, Gabon, Malawi, Mozambique, Sierra Leone, South Africa, Zambia, Zimbabwe)

Coelioxys aurulenta Pasteels, 1968 (South Africa)

Coelioxys bifoliata Pasteels, 1968 (South Africa)

Coelioxys bifoveolata Pasteels, 1968 (Namibia)

Coelioxys binghami Pasteels, 1968 (South Africa)

Coelioxys bruneipes Pasteels, 1968 (South Africa, Zimbabwe)

Coelioxys bulbosa Pasteels, 1985 (Botswana, Burkina Faso, Namibia, Senegal)

Coelioxys caeruleipennis Friese, 1905 (Democratic Republic of the Congo, Kenya, Malawi, South Africa, Uganda, Zimbabwe)

Coelioxys caffra Friese, 1905 (Kenya, Mozambique, Namibia, South Africa, Uganda, Zimbabwe)

Coelioxys calabarensis Pasteels, 1968 (Equatorial Guinea)

Coelioxys capensis Smith, 1854 (Democratic Republic of the Congo, Kenya, South Africa, Zimbabwe)

Coelioxys carinicauda Pasteels, 1968 (South Africa)

Coelioxys cavigena Pasteels, 1968 (Senegal)

Coelioxys cherenensis Friese, 1913 (Eritrea, Ethiopia, Ghana, Sudan, Uganda)

Coelioxys circumscriptus Schulz, 1906 (Lesotho, South Africa. Also in Palaearctic region)

Coelioxys coloratula Cockerell, 1939 (South Africa)

Coelioxys congoensis Friese, 1922 (Congo, Democratic Republic of the Congo, Senegal, Zambia)

Coelioxys coriacea Pasteels, 1968 (Malawi)

Coelioxys coturnix Pérez,, 1884 (Gambia [?], Senegal [?]. Also in Palaearctic region) [Afrotropical material possibly misidentified])

Coelioxys cyanura Cockerell, 1933 (Burundi, Congo, Democratic Republic of the Congo, Ghana, Liberia, Uganda, Zimbabwe)

Coelioxys decipiens Spinola, 1839 (Chad, Democratic Republic of the Congo, Ethiopia, Mauritania, Senegal, South Africa, Sudan. Also in Palaearctic region)

Coelioxys demeter Cockerell, 1939 (South Africa)

Coelioxys desmieri Pasteels, 1968 (Senegal)

Coelioxys difformis Friese, 1905 (South Africa, Zimbabwe)

Coelioxys elegantula Alfken, 1934 (Yemen[?]. Also in Palaearctic region) [African records doubtful]

Coelioxys elongativentris Pasteels, 1968 (Niger)

Coelioxys emarginata onculatus Friese, 1941 (Mozambique)

Coelioxys erythrura Spinola, 1839 (Niger, Senegal)

Coelioxys erythrura rubraxillaris Pasteels, 1968 (Niger)

Coelioxys foveolata Smith, 1854 (Democratic Republic of the Congo, Gabon, Malawi, Mozambique, Somalia, South Africa, Zambia, Zimbabwe)

Coelioxys genalis Pasteels, 1985 (Namibia)

Coelioxys gracillima Pasteels, 1968 (Democratic Republic of the Congo, Zimbabwe)

Coelioxys haemorrhoa Foerster, 1853 (Burundi)

Coelioxys heterozona Cockerell, 1939 (“East Africa”, Ghana, Nigeria, Sudan)

Coelioxys incarinata Friese, 1905 (South Africa, Zimbabwe)

Coelioxys integra Pasteels, 1968 (Cameroun)

Coelioxys intermedia Pasteels, 1968 (South Africa, Sudan)

Coelioxys junodi Friese, 1905 (Congo, South Africa)

Coelioxys katangensis Cockerell, 1932 (Democratic Republic of the Congo)

Coelioxys langi Cockerell, 1935 (Zimbabwe)

Coelioxys lativentris Friese, 1909 (Malawi, South Africa, Zambia, Zimbabwe)

Coelioxys lativentroides Brauns, 1930 (South Africa, Zimbabwe)

Coelioxys longispina Pasteels, 1968 (South Africa)

Coelioxys loricula Smith, 1854 (Gabon, Ghana, Guinea, Kenya, South Africa, Zimbabwe)

Coelioxys luangwana Cockerell, 1939 (Zambia)

Coelioxys lucidicauda Cockerell, 1939 (South Africa)

Coelioxys maculata Friese, 1913 (Democratic Republic of the Congo, Malawi, Mozambique, South Africa, Uganda, “West Africa”)

Coelioxys maculoides Pasteels, (Ivory Coast)

Coelioxys (Liothyrapsis) maculoides Pasteels, 1968a: 101, 132, male holotype (MNHN) Ivory Coast)

Liothyrapsis (Torridapis) maculoides (Pasteels): Pasteels 1977b: 195)

Coelioxys (Torridapis) maculoides Pasteels: Michener 2000: 530 [by implication])

Coelioxys madagascariensis Benoist, 1955 (Madagascar)

Coelioxys magretti Friese, 1915 (Ethiopia)

Coelioxys marchalli Pasteels, 1968 (Zimbabwe)

Coelioxys nasuta Friese, 1905 (Democratic Republic of the Congo, Kenya, Malawi, Mali, Niger, Senegal, South Africa, Zambia, Zimbabwe)

Coelioxys natalensis Cockerell, 1920 (Democratic Republic of the Congo, “East Africa”, Gabon, Kenya, Malawi, South Africa, Uganda, Zambia, Zimbabwe)

Coelioxys neli Cockerell, 1933 (South Africa)

Coelioxys nigripes Vachal, 1903 (Gabon, Nigeria, Sierra Leone)

Coelioxys nigrura Pasteels, 1977 (Senegal)

Coelioxys nitidicauda Pasteels, 1968 (South Africa)

Coelioxys nivosa Pasteels, 1968 (Somalia)

Coelioxys odin Strand, 1912 (Cameroun, Democratic Republic of the Congo, Ethiopia, Equatorial Guinea, South Africa, Tanzania, Uganda)

Coelioxys paludicola Strand, 1911 (South Africa)

Coelioxys penetratrix Smith, 1879 (Malawi, South Africa)

Coelioxys piligena Pasteels, 1968 (Ethiopia, Malawi)

Coelioxys planidens Friese, 1905 (Democratic Republic of the Congo, Malawi, Mozambique, Senegal, South Africa)

Coelioxys postponenda Schulz, 1906 (Guinea, Liberia, Malawi, Mali, Senegal, South Africa)

Coelioxys raffrayi Dusmet y Alonso, 1912 (Distribution unknown)

Coelioxys recusata Schulz, 1906 (Democratic Republic of the Congo, Kenya, South Africa, Uganda)

Coelioxys reticulata Pasteels, 1968 (Senegal)

Coelioxys rotundiscutum) (Pasteels, 1977) (Senegal)

Coelioxys rubella Pasteels, 1968 (Senegal)

Coelioxys rufispina Walker, 1871 (South Africa, Sudan, Zimbabwe. Also in Palaearctic region)

Coelioxys scioensis Gribodo, 1879 (Eritrea, Ethiopia, Guinea, Kenya, Malawi, Nigeria, Sierra Leone, Tanzania, Uganda)

Coelioxys scutellotuberculata Pasteels, 1968 (Niger)

Coelioxys seminitida Pasteels, 1985 (South Africa)

Coelioxys setosa Friese, 1905 (Angola, Malawi, Mozambique, Senegal, South Africa, Tanzania, Zambia)

Coelioxys somalica Friese, 1922 (Namibia, Somalia, Zambia, Zimbabwe)

Coelioxys somalina Magretti, 1895 (Malawi, Zambia, Zimbabwe)

Coelioxys spissicauda Pasteels, 1968 (South Africa)

Coelioxys squamosa Friese, 1922 (Namibia, South Africa, Zimbabwe)

Coelioxys squamosissima Pasteels, 1968 (South Africa)

Coelioxys squamosoides Pasteels, 1977 (Zambia)

Coelioxys squamatula Pasteels, 1968 (Rwanda)

Coelioxys sudanensis Cockerell, 1933 (Sudan)

Coelioxys subdentata Smith, 1854 (Democratic Republic of the Congo, South Africa)

Coelioxys subnitens Pasteels, 1977 (Senegal)

Coelioxys tenebrosa Pasteels, 1968 (Zimbabwe)

Coelioxys tenebrosoides Pasteels, 1968 (South Africa, Zambia)

Coelioxys togoensis Pasteels, 1968 (Togo)

Coelioxys torrida Smith, 1854 (Angola, Botswana, Democratic Republic of the Congo, Kenya, Malawi, Mozambique, Sierra Leone, South Africa, Tanzania, Togo, Uganda, Zimbabwe)

Coelioxys torridula Pasteels, 1968 (Mauritius)

Coelioxys truncaticauda Cockerell, 1932 (Namibia, South Africa)

Coelioxys ultima Pasteels, 1968 (Uganda)

Coelioxys umbripennis Friese, 1922 (Botswana, Democratic Republic of the Congo, Ghana, Guinea, Kenya, Malawi, Senegal, South Africa, Uganda, Zambia, Zimbabwe)

Coelioxys unicula Cockerell, 1939 (South Africa, Zimbabwe)

Coelioxys verticalis Smith, 1854 (Democratic Republic of the Congo, Liberia, Sierra Leone, South Africa, Uganda)

Comments	The genus Coelioxys is split into four subgenera (text verbatim from Eardley et al. 2010): *Subgenus Coelioxys (Allocoelioxys)* Tkalcu This subgenus is widespread in the Old World and occurs throughout Africa. Pasteels (1977) recorded 21 African species. Subgenus Coelioxys (Coelioxys) Latreille This subgenus occurs throughout Africa, and is apparently cosmopolitan. Pasteels (1977) recorded 31 African species. Subgenus Coelioxys (Liothyrapis) Cockerell This subgenus occurs throughout Africa and is widely distributed in the Old World. Pasteels (1968) recorded 26 species from sub-Saharan Africa. Subgenus Coelioxys (Torridapis) Pasteels** This subgenus is widespread in Africa and it is also widespread in southern Asia, the Pacific Islands and Australia. There are seven African species.
Distribution	Worldwide.
Biology	Coelioxys species are cleptoparasitic, mostly parasitizing Megachile species (Eardley et al., 2010).
References	Brothers D.J. 1999. Phylogeny and evolution of wasps, ants and bees (Hymenoptera, Chrysidoidea, Vespoidea and Apoidea) Zoologica Scripta 28: 233–250. Eardley, C & Urban, R. 2010. Catalogue of Afrotropical bees (Hymenoptera: Apoidea: Apiformes). Zootaxa 2455: 1-548. Eardley, C, Finnamore, A.T. & Michener, C.D. 1993. Superfamily Apoidea (pp. 279-357). In GOULET, H. & HUBER, J. (eds). Hymenoptera of the World: an identification guide to families. Research Branch, Agriculture Canada, Ottawa, Canada, 668 pp. Eardley C.,Kuhlmann M., Pauly A. 2010. The Bee Genera and Subgenera of sub-Saharan Africa. Abc Taxa vol 7: i-vi, 138 pp. Michener, C.D. 2000. The Bees of the World. Johns Hopkins University Press. 953 pp. Pasteels, J.J. 1968. Révision des Megachilidae (Hymenoptera Apoidea) de L'Afrique Noire. II. Le Genre Coelioxys. Musée Royal de L'Afrique Centrale Tervuren, Belgique, Annales, Serie In-8- Sciences Zoologiques 167: 1-139. Pasteels, J.J. 1977. Les Megachilini parasites (Coelioxys s.l.) d'Afrique noire. Revue de Zoologie Africaine 91: 1-197.
Credits	Photographs © Dave West (Randburg).

Coelioxys Latreille

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