Bethylidae

Classification of afrotropical Bethylid wasps

(Life: Kingdom: Metazoa (animals); Phylum: Arthropoda; Class: Hexapoda; Order: Hymenoptera; Superfamily: Chrysidoidea; Family: Bethylidae)

Subfamily BETHYLINAE
- Genus Goniozus Föster, 1856
  - Goniozus etiellae Risbec, 1955
  - Goniozus garouae (Risbec, 1955) (Senegal)
  - Goniozus kabisus (Benoit, 1957) (Democratic Republic of Congo)
  - Goniozus leuconeura (Kieffer, 1914) (Namibia)
  - Goniozus procerae Risbec, 1956 (Cameroon, Senegal)
  - Goniozus rostratus Kieffer, 1905 (Madagascar)
  - Goniozus rutshurus (Benoit, 1957) (Democratic Republic of Congo)
  - Goniozus simillis Fouts, 1934 (Somalia)
- Genus Trissomalus Kieffer, 1905
  - Trissomalus cameroni Kieffer, 1914 (South Africa)
Subfamily EPYRINAE
- Genus Calyoza Westwood, 1837
  - Calyoza staphylinoides Hope, 1837 (South Africa)
  - Calyoza ashmeadi Ebderlein, 1901 (Cameroon)
- Genus Calyozina Enderlein, 1912
  - Calyozina caperata Krombein, 1990 (Botswana)
- Genus Epyris Westwood, 1832
  - Epyris algoae Kieffer, 1911 (South Africa)
  - Epyris algoaensis Kieffer, 1911 (South Africa)
  - Epyris allaudi Kieffer, 1905 (Democratic Republic of Congo, Madagascar)
  - Epyris amabilis Saussure, 1892 (Democratic Republic of Congo)
  - Epyris bicolor Fouts, 1934 (Somalia)
  - Epyris caffrarius Kieffer, 1911 (South Africa)
  - Epyris congolensis Benoit, 1957 (Democratic Republic of Congo)
  - Epyris ebneri Kieffer, 1921 (Sudan, Democratic Republic of Congo)
  - Epyris fallax Benoit, 1957 (Democratic Republic of Congo)
  - Epyris fuelleborni Kieffer, 1912 (Malawi)
  - Epyris geniculatus Kieffer, 1904 (Democratic Republic of Congo, Eritrea)
  - Epyris hottentottus Turner, 1928 (South Africa)
  - Epyris latipes Kieffer, 1913 (Democratic Republic of Congo, Eritrea)
  - Epyris levissimus Turner, 1928 (South Africa)
  - Epyris lippensi Benoit, 1957 (Democratic Republic of Congo)
  - Epyris michaelseni Kieffer, 1914 (Namibia)
  - Epyris muiri Turner and Waterston, 1917 (South Africa)
  - Epyris naticola Kieffer, 1913 (South Africa)
  - Epyris perversus Benoit, 1957 (Democratic Republic of Congo)
  - Epyris plurilineatus Turner, 1928 (Democratic Republic of Congo, South Africa, Zimababwe)
  - Epyris ruficaudus (Kieffer, 1913) (South Africa)
  - Epyris silverstrii Kieffer, 1913 (Ghana)
  - Epyris subalatus Turner, 1928 (South Africa)
  - Epyris torridus Kieffer, 1912 (Tanzania)
  - Epyris windhukiensis Kieffer, 1914 (Namibia)
  - Epyris wittei Benoit, 1957 (Democratic Republic of Congo)
- Genus Glenosema Kieffer, 1905
  - Glenosema viduatus Turner, 1928 (South Africa)
- Genus Holepyris Kieffer, 1905
  - Holepyris africanus Kieffer, 1904 (Eritrea)
  - Holepyris algoaensis Kieffer, 1911 (South Africa)
  - Holepyris capicola Kieffer, 1911 (South Africa)
  - Holepyris cerostrotoideus Benoit, 1957 (Democratic Republic of Congo)
  - Holepyris cerostrotus Benoit, 1957 (Democratic Republic of Congo)
  - Holepyris clytus Benoit, 1957 (Democratic Republic of Congo)
  - Holepyris natalensis Kieffer, 1913 (South Africa)
  - Holepyris plebejus Benoit, 1957 (Democratic Republic of Congo)
  - Holepyris pondo Turner, 1928 (South Africa)
  - Holepyris rwindius Benoit, 1957 (Democratic Republic of Congo)
  - Holepyris semirruber Kieffer, 1911 (South Africa)
  - Holepyris sylvaticus (Turner, 1928) (South Africa)
- Genus Homoglenus Kieffer, 1904
  - Homoglenus quadripartitus Benoit, 1957 (Democratic Republic of Congo)
  - Homoglenus sanctus Turner, 1928 (South Africa)
- Genus Melanepyris Kieffer, 1913
  - Melanopyris imicola Kieffer, 1913 (Kenya)
- Genus Messoria Meunier, 1916
  - Messoria copalina Meunier, 1916 (Kenya)
- Genus Neurepyris Kieffer, 1905
  - Neurepyris rufiventer Kieffer, 1913 (Eritrea)
- Genus Planepyris Kieffer, 1905
  - Planepyris appendiculatus Kieffer, 1913 (Eritrea)
- Genus Plastanoxus Kieffer, 1905
  - Plastanoxus westwoodi (Kieffer, 1914) (Australia, Columbia, England, Finland, Italy, U.S.A)
- Genus Pristepyris Kieffer, 1905
  - Pristepyris flavicornis Kieffer, 1912 (Cameroon)
  - Pristepyris levicollis Kieffer, 1905 (Madagascar)
- Genus Prolaelius Kieffer, 1905
  - Prolaelius firmipennis Cameron, 1905 (South Africa)
  - Prolaelius glossinae (Turner & Waterston, 1916) (Malawi)
- Genus Prorops Waterston, 1923
  - Prorops nasuta Waterson, 1923 (Uganda)
- Genus Pseudocalyoza Turner, 1915
  - Pseudocalyoza subramosa Turner, 1915 (South Africa)
- Genus Rhabdepyris Kieffer, 1904
  - Rhabdepyris capensis Kieffer, 1911 (South Africa)
  - Rhabdepyris dilaticeps Benoit, 1957 (Democratic Republic of Congo)
- Genus Sclerodermus Latreille, 1809
  - Sclerodermus cadavericus Benoit, 1957 (Democratic Republic of Congo, Kenya, Uganda)
  - Sclerodemus wollastoni Westwood, 1881 (Sierra Leone, St. Helene)
- Genus Trachepyris Kieffer, 1905
  - Trachepyris breviscapus (Kieffer, 1904) (Cape Verde Islands, Democratic Republic of Congo, Guinea, Bissau, Rwanda, South Africa)
  - Trachepyris ceresensis (Turner, 1928) (Democratic Republic of Congo, South Africa)
  - Trachepyris crenaticollis (Kieffer, 1912) (Malawi)
  - Trachepyris hildebrandti (Kieffer, 1912) (Tanzania)
  - Trachepyris semiserratus (Kieffer, 1911) (South Africa)
  - Trachepyris serricollis (Westwood, 1874) (South Africa)
  - Trachepyris spiniscapus (Kieffer, 1904) (Eritrea)
- Genus Trissepyris Kieffer, 1905
  - Trissepyris ruficeps Kieffer, 1905 (Democratic Republic of Congo)

Subfamily MESITIINAE Berland, 1928
- Genus Heterocoelia Dhlbom, 1854
  - Heterocoelia boharti Moczar, 1971 (South Africa)
  - Heterocoelia brauns (Kieffer, 1911) (South Africa)
  - Heterocoelia fischeri Moczar, 1971 (South Africa)
  - Heterocoelia fortidens (Kieffer, 1913) (South Africa)
  - Heterocoelia paenepunctata (Benoit, 1968) (Democratic Republic of Congo)
  - Heterocoelia rhodesica (Moczar, 1971 (Zimbabwe)
  - Heterocoelia tsunekii (Moczar, 1971) (Ghana)
- Genus Metrionotus Moczar, 1970
  - Metrionotus africanus Moczar, 1970 (South Africa)
  - Metrionotus alutaceus (Benoit, 1968) (South Africa)
  - Metrionotus bachmaieri Moczar, 1970 (South Africa)
  - Metrionotus bouceki (Moczar, 1971) (South Africa)
  - Metrionotus brevispinosus (Benoit, 1968) (Botswana, Namibia, South Africa)
  - Metrionotus carbonarius Moczar, 1970 (Hungary, South Africa)
  - Metrionotus laterinotus Moczar, 1970 (Namibia)
  - Metrionotus minor (Benoit, 1968) (South Africa)
  - Metrionotus minutissimus (Moczar, 1971) (South Africa)
  - Metrionotus mocsaryi Moczar, 1970 (South Africa)
  - Metrionotus nigropiceus (Moczar, 1971) (South Africa)
  - Metrionotus pappi Moczar, 1970 (South Africa)
  - Metrionotus szelenyii Moczar, 1970 (Namibia, South Africa, Yemen, )
  - Metrionotus turneri (Benoit, 1968) (South Africa)
  - Metrionotus wolfi (Moczar, 1970 (South Africa)
  - Metrionotus yarrowi Moczar, 1970 (South Africa)
  - Metrionotus zuluensis (Moczar, 1971) (South Africa)
  - Metrionotus parvulus (Kieffer, 1906) (Italy, Tanzania)
- Genus Parvoculus Moczar, 1970
  - Parvoculus myrmecophilus Moczar, 1970 (Democratic Republic of Congo)
- Genus Pilomesitius Moczar, 1970
  - Pilomesitius madagascarensis Moczar, 1970 (Madagascar)
- Genus Pycnomesitius Moczar, 1971
  - Pycnomesitius brevialatus (Benoit, 1968) (South Africa)
  - Pycnomesitius densepunctatus Moczar, 1971 (South Africa)
  - Pycnomesitius peringueyi (Kieffer, 1913) (South Africa)
- Genus Sulcomesitius Moczar, 1970
  - Sulcomesitius africanus Moczar, 1970 (South Africa)
  - Sulcomesitius alluadi (Kieffer, 1913) (Democratic Republic of Congo, Kenya)
  - Sulcomesitius arushai (Moczar, 1969) (Tanzania)
  - Sulcomesitius benoiti Moczar, 1970 (South Africa)
  - Sulcomesitius brevidens (Kieffer, 1913) (South Africa)
  - Sulcomesitius bridwell (Nagy, 1968) (Democratic Republic of Congo, Nigeria)
  - Sulcomesitius capensis (Kieffer, 1911) (South Africa, Zambia, Zimbabwe)
  - Sulcomesitius consimilis Moczar, 1970 (South Africa)
  - Sulcomesitius erdoesi Moczar, 1970 (South Africa)
  - Sulcomesitius grahamensis Moczar, 1984 (South Africa)
  - Sulcomesitius hilarius Nagy, 1972 (Ethiopia)
  - Sulcomesitius illaevigatus (Benoit, 1968) (Democratic Republic of Congo, Kenya)
  - Sulcomesitius katonai (Moczar, 1970) (Tanzania)
  - Sulcomesitius linsenmaieri Moczar, 1970 (Senegal)
  - Sulcomesitius longispinus Moczar, 1970 (South Africa)
  - Sulcomesitius madagascarensis Moczar, 1970 (Madagascar)
  - Sulcomesitius nasutus (Benoit, 1968) (South Africa)
  - Sulcomesitius okahandjaensis Moczar, 1977 (Namibia)
  - Sulcomesitius pondo (Benoit, 1968) (India, South Africa)
  - Sulcomesitius rufus (Moczar, 1969) (South Africa)
  - Sulcomesitius saintjohni Moczar, 1977 (South Africa)
  - Sulcomesitius schoutedeni (Benoit, 1968) (West Democratic Republic of Congo, Tanzania)
  - Sulcomesitius somalicus (Masi, 1936) (Somalia)
  - Sulcomesitius townesianus Moczar, 1983 (Uganda)
  - Sulcomesitius turneri Moczar, 1970 (South Africa)
  - Sulcomesitius zambiensis Moczar, 1984 (Zambia)
Subfamily PRISTOCERINAE
- Genus Afgoiogfa Argaman, 1988
  - Afgoiogfa olmiana Argaman, 1988 (Somalia)
- Genus Afrocera Benoit, 1983
  - Afrocera bamboutoana Benoit, 1983 (Cameroon)
- Genus Apenesia Westwood, 1874
  - Apenesia acuta Benoit, 1957 (Democratic Republic of Congo)
  - Apenesia conradti Kieffer, 1912 (Cameroon)
  - Apenesia forchammeri Krombein, 1989 (Botswana)
  - Apenesia levis Kieffer, 1904 (West Africa)
  - Apenesia nigra Kieffer, 1904 (West Africa)
  - Apenesia nyama (Benoit, 1957) (Democratic Republic of Congo)
  - Apenesia nyamuragira (Benoit, 1957) (Democratic Republic of Congo)
  - Apenesia pseudosuzannae (Benoit, 1957) (Democratic Republic of Congo)
  - Apenesia punctulata Krombein, 1904 (Cameroon)
  - Apenesia remota (Kieffer, 1912) (Seychelles)
  - Apenesia sjostedti (Tulgren, 1904) (Cameroon)
  - Apenesia suzannae (Benoit, 1957) (Democratic Republic of Congo)
  - Apenesia tridentata Kieffer, 1912 (Cameroon)
  - Apenesia triloba (Benoit, 1957) (Democratic Republic of Congo, South Africa)
- Genus Apristocera Kieffer, 1914
  - Apristocera natalensis (Kieffer, 1911) (South Africa)
- Genus Dicrogenium Stadelmann, 1894
  - Dicrogenium aethiopicum Benoit, 1963 (Ethiopia)
  - Dicrogenium africanum (Kieffer, 1911) (Zimbabwe)
  - Dicrogenium alberti Benoit, 1957 (Democratic Republic of Congo)
  - Dicrogenium armaticeps (Turner, 1915) (Ethiopia)
  - Dicrogenium auratum Benoit, 1963 (Democratic Republic of Congo, Tanzania)
  - Dicrogenium bequaerti Benoit, 1963 (Democratic Republic of Congo)
  - Dicrogenium bicarinatum Benoit, 1963 (Zambia, Zimbabwe)
  - Dicrogenium centrale Benoit, 1956 (Burundi, Zambia)
  - Dicrogenium chirindaense Benoit, 1963 (Zimbabwe)
  - Dicrogenium collare Benoit, 1963 (Burundi, Zambia)
  - Dicrogenium conradti Stadelamann, 1894 (Cameroon, Namibia, Togo)
  - Dicrogenium elongatum Benoit, 1963 (Democratic Republic of Congo)
  - Dicrogenium giganteum Arle, 1930 (Gabon)
  - Dicrogenium liberiense Benoit, 1963 (Liberia)
  - Dicrogenium maximum Turner, 1917 (Sierra Leone)
  - Dicrogenium mufungwaense Benoit, 1963 (Democratic Republic of Congo)
  - Dicrogenium rosmarum (Stadelmann, 1892) (Cameroon)
  - Dicrogenium schoutedeni Benoit, 1963 (Democratic Republic of Congo)
  - Dicrogenium spinatum Benoit, 1963 (Democratic Republic of Congo, Malawi)
  - Dicrogenium spiniceps Masi, 1939 (Ethiopia)
  - Dicrogenium thermophilum Benoit, 1957 (Democratic Republic of Congo)
  - Dicrogenium urundiense Benoit, 1956 (Burundi)
- Genus Diepyris Benoit, 1957
  - Diepyris abdominalis Turner, 1915 (Democratic Republic of Congo, Uganda)
  - Diepyris brunneus Benoit, 1957 (Democratic Republic of Congo)
  - Diepyris congoensis Benoit, 1982 (Democratic Republic of Congo)
  - Diepyris lembanus Benoit, 1982 (Democratic Republic of Congo)
  - Diepyris tanganus Benoit, 1982 (Tanzania)
  - Diepyris uniforms Benoit, 1982 (Democratic Republic of Congo)
- Genus Dissomphalus Ashmead, 1893
  - Dissomphalus adscitus Benoit, 1956 (Democratic Republic of Congo, Rwanda)
  - Dissomphalus alberti Benoit, 1957 (Democratic Republic of Congo)
  - Dissomphalus clibanarius Benoit, 1957 (Democratic Republic of Congo)
  - Dissomphalus crucialis Benoit, 1957 (Democratic Republic of Congo)
  - Dissomphalus excisicrus Kieffer, 1912 (Seychelles)
  - Dissomphalus globiceps Benoit, 1957 (Democratic Republic of Congo)
  - Dissomphalus indeceptibilis Benoit, 1957 (Democratic Republic of Congo)
  - Dissomphalus nicaeus Benoit, 1956 (Democratic Republic of Congo, Rwanda)
  - Dissomphalus orophilus Benoit, 1956 (Democratic Republic of Congo, Rwanda)
  - Dissomphalus prophylax Benoit, 1957 (Democratic Republic of Congo)
  - Dissomphalus rwindianus Benoit, 1957 (Democratic Republic of Congo)
  - Dissomphalus saxatilis Kieffer, 1912 (Seychelles)
  - Dissomphalus species
- Genus Kathepyris Kieffer, 1907
  - Kathepyris basutoensis Benoit, 1982 (Lesotho)
  - Kathepyris katangensis Benoit, 1982 (Democratic Republic of Congo)
  - Kathepyris nyassica Kieffer, 1907 (Malawi)
  - Kathepyris uelensis Benoit, 1982 (Democratic Republic of Congo)
- Genus Neodicrogenium Benoit, 1957
  - Neodicrogenium beqaerti Benoit, 1963 (Democratic Republic of Congo)
  - Neodicrogenium commune (Turner, 1915) (Malawi)
  - Neodicrogenium sampwense Benoit, 1963 (Democratic Republic of Congo)
  - Neodicrogenium spina Benoit, 1957 (Democratic Republic of Congo)
  - Neodicrogenium superbum Benoit, 1963 (Democratic Republic of Congo)
  - Neodicrogenium tuberculatum (Turner, 1915) (Malawi)
- Genus Neusakosia Benoit, 1981
  - Neusakosia princeps Benoit, 1981 (Namibia)
  - Neusakosia schoutedeni Benoit, 1981 (Namibia)
- Genus Parapenesia Kieffer, 1910
  - Parapenesia unicolor Kieffer, 1910 (South Africa)
- Genus Parascleroderma Kieffer, 1904
  - Parascleroderma nigra Brues, 1910 (South Africa)
  - Parascleroderma seychelles Kieffer, 1912 (South Africa)
- Genus Pristocera Klug, 1808
  - Pristocera atopogamia (Turner, 1915) (Democratic Republic of Congo, Malawi, Namibia, Zimbabwe)
  - Pristocera axitiosa Benoit, 1957 (Democratic Republic of Congo)
  - Pristocera basilewskyi Benoit, 1956 (Democratic Republic of Congo)
  - Pristocera bequaerti Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera camerunsis Benoit, 1963 (Cameroon)
  - Pristocera capensis Benoit, 1963 (South Africa)
  - Pristocera cara Benoit, 1957 (Democratic Republic of Congo)
  - Pristocera chillaloensis Benoit, 1963 (Ethiopia)
  - Pristocera communis (Turner, 1915) (South Africa)
  - Pristocera congoensis Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera decellei Benoit, 1963 (Ivory Coast)
  - Pristocera decemdentata Enderlein, 1901 (Cameroon, Democratic Republic of Congo, Togo)
  - Pristocera distincta Benoit, 1963 (South Africa)
  - Pristocera draconum Turner, 1928 (South Africa)
  - Pristocera elongata Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera erythrura Kieffer, 1904 (Somalia)
  - Pristocera fallax Benoit, 1963 (South Africa)
  - Pristocera fellimontana Benoit, 1963 (South Africa)
  - Pristocera ferruginea Bischoff, 1915 (Tanzania)
  - Pristocera fossulata Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera gaullei Kieffer, 1908 (Benin, Guinea, South Africa)
  - Pristocera hirsutula Turner, 1928 (Lesotho, South Africa)
  - Pristocera humilis Benoit, 1963 (South Africa)
  - Pristocera incerta (Turner, 1915) (Malawi)
  - Pristocera jacoti Benoit, 1963 (Lesotho, South Africa)
  - Pristocera juncta Turner, 1928 (South Africa)
  - Pristocera katangensis Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera kenyaensis Benoit, 1963 (Kenya)
  - Pristocera kiefferi Benoit, 1963 (South Africa)
  - Pristocera kwangoa Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera laticornis Kieffer, 1913 (Malawi, Zimbabwe)
  - Pristocera luminosa Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera meridionalis Benoit, 1963 (South Africa)
  - Pristocera natalensis Benoit, 1963 (South Africa)
  - Pristocera neavei Turner and Waterston, 1917 (Mozambique)
  - Pristocera nigerrima Benoit, 1957 (Democratic Republic of Congo, Rwanda)
  - Pristocera nigrita Kieffer, 1905 (Democratic Republic of Congo, Madagascar)
  - Pristocera obliterata Kieffer, 1913 (Democratic Republic of Congo, Kenya, Rwanda)
  - Pristocera oriphila Turner, 1928 (South Africa)
  - Pristocera pagana Benoit, 1957 (Democratic Republic of Congo)
  - Pristocera pondoensis Benoit, 1963 (South Africa)
  - Pristocera regina Benoit, 1957 (Democratic Republic of Congo, Rwanda)
  - Pristocera rhodesiae Turner, 1928 (Zimbabwe)
  - Pristocera rufabdominalis Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera ruficaudata Westwood, 1874 (Madagascar, South Africa)
  - Pristocera rugosa Enderlein, 1901 (Democratic Republic of Congo, Togo)
  - Pristocera silvatica Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera subrufescens Turner and Waterston, 1917 (South Africa)
  - Pristocera subviolacea Enderlein, 1901 (Cameroon, Dahomey)
  - Pristocera sulcata Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera termitophila Benoit, 1963 (South Africa)
  - Pristocera tuberculata (Turner, 1915) (Malawi)
  - Pristocera turneri Benoit, 1963 (South Africa)
  - Pristocera uelensis Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera vandenbrandei Benoit, 1963 (Democratic Republic of Congo)
  - Pristocera vumbua Benoit, 1963 (Zimbabwe)
- Genus Prosapenesia Kieffer, 1910
  - Prosapenesia bilobata Benoit, 1981 (Namibia)
  - Prosapenesia lacteipennis Kieffer, 1910 (Namibia)
- Genus Protisobrachium Benoit, 1957
  - Protisbrachium gracile Benoit 1957 (Democratic Republic of Congo)
  - Protisbrachium rwindicum Benoit, 1957 (Democratic Republic of Congo)
- Genus Pseudisobrachium Kieffer, 1904
  - Pseudisobrachium babaeculum Benoit, 1957 (Democratic Republic of Congo)
  - Pseudisobrachium babaeculuscum (Benoit, 1957) (Democratic Republic of Congo)
  - Pseudisobrachium cruciferum (Kieffer, 1912) (Seychelles)
  - Pseudisobrachium kiefferi Benoit, 1957 (Democratic Republic of Congo)
  - Pseudisobrachium matticum Benoit, 1957 (Democratic Republic of Congo)
  - Pseudisobrachium monticola Kieffer, 1913 (Kenya)
  - Pseudisobrachium pallidicorne Kieffer, 1910 (Algeria, Democratic Republic of Congo, Gabon, Israel, Jordon, Morocco, Nigeria, Tunisia)
  - Pseudisobrachium pauculihirtum Benoit, 1957 (Democratic Republic of Congo)
  - Pseudisobrachium transkeiensis (Turner, 1928) (South Africa)
- Genus Trichiscus Benoit, 1956
  - Trichiscus suzannae Benoit, 1986 (Democratic Republic of Congo)
  - Trichiscus wittei Benoit, 1956 (Democratic Republic of Congo, Rwanda)
- Genus Usakosia Kieffer, 1914
  - Usakosia albipennis Kieffer, 1914 (Namibia)